Phase contrast image of biomass from the Tasmanian winery WWTP and corresponding composite FISH image: EUBmix (FLUOS) alone = green; EUBmix (FLUOS) + AMAR839 (CY3) = yellow. Scale bar = 10 microns. - Source:1Aerobic heterotroph
Tetrad forming organisms orignally believed to be GAOs. However pure cultures were shown to be unable to assimilate carbon under anaerobic conditions 2. Aerobic heterotrophs, which have a high capacity for PHA storage 2. No information on their in situ physiology in activated sludge is availabe.
Has been associated with non-filamentous viscous bulking in industrial plants 3.
 McIlroy SJ (2010) Phylogenetic Diversity and Ecophysiology of Alphaproteobacterial Glycogen Accumulating Organisms in Enhanced Biological Phosphorus Removal Activated Sludge Systems. PhD Thesis La Trobe University. - Mcilroy -
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 McIlroy, Speirs, Tucci, Seviour (2011): In situ profiling of microbial communities in full-scale aerobic sequencing batch reactors treating winery waste in australia. Environ. Sci. Technol. 45 (20): 8794-803. doi:10.1021/es2018576
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